Age Differences in Flight Feathers
There are often substantial differences between the remiges and rectrices of adults and juveniles of the same species. Because all juvenile feathers are grown at once—a tremendous energy burden to the developing bird—they are softer and of poorer quality than the equivalent feathers of adults, which are moulted over a longer period of time (as long as several years in some cases). As a result, they wear more quickly.
As feathers grow at variable rates, these variations lead to visible dark and light bands in the fully formed feather. These so-called growth bars and their widths have been used to determine the daily nutritional status of birds. Each light and dark bar correspond to around 24 hours and the use of this technique has been called ptilochronology (analogous to dendrochronology).
In general, juveniles have feathers which are narrower and more sharply pointed at the tip. This can be particularly visible when the bird is in flight, especially in the case of raptors. The trailing edge of the wing of a juvenile bird can appear almost serrated, due to the feathers' sharp tips, while that of an older bird will be straighter-edged. The flight feathers of a juvenile bird will also be uniform in length, since they all grew at the same time. Those of adults will be of various lengths and levels of wear, since each is moulted at a different time.
The flight feathers of adults and juveniles can differ considerably in length, particularly among the raptors. Overall, juveniles tend to have slightly longer rectrices, and shorter, broader wings (with shorter outer primaries, and longer inner primaries and secondaries) than do adults of the same species. However, there are many exceptions. In longer-tailed species, such as Swallow-tailed Kite, Secretary Bird and European Honey Buzzard, for example, juveniles have shorter rectrices than adults do. Juveniles of some Buteo buzzards have narrower wings than adults do, while those of large juvenile falcons are longer. It is theorized that the differences help young birds compensate for their inexperience, weaker flight muscles and poorer flying ability.
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