Early Life
Guthrie received his doctorate from the University of Minnesota, although his education took a circuitous route, as he eventually earned six degrees in six years, including both a medical doctorate and a doctor of philosophy. While in school, Guthrie married Margaret, a fellow student, and they eventually had six children together. His early research into bacterial inhibition assays came while he was employed by the Staten Island Public Health Hospital, testing antibiotic sensitivity.
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