Physiological Basis
The physiological basis for color constancy is thought to involve specialized neurons in the primary visual cortex that compute local ratios of cone activity, which is the same calculation that Land's retinex algorithm uses to achieve color constancy. These specialized cells are called double-opponent cells because they compute both color opponency and spatial opponency. Double-opponent cells were first described by Nigel Daw in the goldfish retina. There was considerable debate about the existence of these cells in the primate visual system; their existence was eventually proven using reverse-correlation receptive field mapping and special stimuli that selectively activate single cone classes at a time, so-called "cone-isolating" stimuli.
Color constancy works only if the incident illumination contains a range of wavelengths. The different cone cells of the eye register different ranges of wavelengths of the light reflected by every object in the scene. From this information, the visual system attempts to determine the approximate composition of the illuminating light. This illumination is then discounted in order to obtain the object's "true color" or reflectance: the wavelengths of light the object reflects. This reflectance then largely determines the perceived color.
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