Hardiness
Typically, overwintering organisms can either avoid freezing or tolerate freezing, but Excechia nugatoria can do both. For E. nugatoria, the production of noncolligative antifreeze proteins (NAPs) protect the head and thorax from freezing, but they do not protect the abdomen.
Freezing of the head and thorax in other insects tends to have adverse effects on neural tissue, so it is not surprising that these protective mechanisms have been observed in certain species, but E. nugatoria is the only insect known to semi-freeze through the winter, which may be an evolutionary advantage according to Sformo: By allowing the abdomen to freeze, evaporative water loss is reduced over the course of the winter.
Read more about this topic: Fungus Gnat