Academic Career
He studied botany, physics and pharmacology at the University of Göttingen under Friedrich Wöhler (1800-1882), William Eduard Weber (1804-1891) and Wilhelm Rudolph Fittig (1835-1910). Afterwards, he furthered his education at the Universities of Marburg and Berlin. At Berlin, he studied under Alexander Braun (1805-1877) and was an assistant to Nathanael Pringsheim (1823-1894). Later on, he served as an assistant to Julius von Sachs (1832-1897) at Würzburg,
In 1873 he was appointed professor of pharmacology and botany at the University of Bonn, followed by professorships at the Universities of Basel (from 1877) and Tübingen (from 1878), where he also served as director of the Botanischer Garten der Universität Tübingen. In 1887 he became a professor at the University of Leipzig and director of its botanical garden.
He was elected a member of the Royal Swedish Academy of Sciences in 1897.
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