Vipera Ammodytes - Taxonomy

Taxonomy

This species was originally described by Carl Linnaeus in Systema Naturae in 1758. Subsequently, George Albert Boulenger described a number of subspecies in the early 20th century that are still mostly recognized today. However, there are many alternative taxonomies. One additional subspecies that may be encountered in literature is V. a. ruffoi (Bruno, 1968), found in the Alpine region of Italy. However, many consider both V. a. ruffoi and V. a. gregorwalineri to be synonymous with V. a. ammodytes and V. a. transcaucasiana to be a separate species.

Subspecies Taxon author Common name Geographic range
V. a. ammodytes (Linnaeus, 1758) Western sand viper Austria (Styria, Carinthia), north Italy, Slovenia, Croatia, Bosnia-Herzegovina, Serbia, Montenegro, Macedonia, Albania, south-west Romania, north-west Bulgaria
V. a. gregorwallneri Sochurek, 1974 Austria, Slovenia, Croatia, Bosnia-Herzegovina, Serbia, Montenegro, Macedonia
V. a. meridionalis Boulenger, 1903 Eastern sand viper Greece (incl. Corfu and other islands), Turkish Thrace
V. a. montandoni Boulenger, 1904 Transdanubian sand viper Bulgaria, south Romania
V. a. transcaucasiana Boulenger, 1913 Transcaucasian sand viper Georgia, north Turkish Anatolia

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