Arthropods
Insects and some more advanced spiders have spiracles on their exoskeletons to allow air to enter the trachea. In the respiratory system of insects, the tracheal tubes primarily deliver oxygen directly into the animals' tissues. The spiracles can be opened and closed in an efficient manner to reduce water loss. This is done by contracting closer muscles surrounding the spiracle. In order to open, the muscle relaxes. The closer muscle is controlled by the central nervous system but can also react to localized chemical stimuli. Several aquatic insects have similar or alternative closing methods to prevent water from entering the trachea. Spiracles may also be surrounded by hairs to minimize bulk air movement around the opening, and thus minimize water loss.
Read more about this topic: Spiracle