Pelagornithidae - Taxonomy, Systematics and Evolution

Taxonomy, Systematics and Evolution

The name "pseudodontorns" refers to the genus Pseudodontornis, which for some time served as the family's namesake. However, the presently used name Pelagornithidae pre-dates Pseudodontornithidae, and thus modern authors generally prefer "pelagornithids" over "pseudodontorns". The latter name is generally found in mid-20th century literature however.

Historically, the disparate bones of the pseudotooth birds were spread across six groups: a number of genera described from leg bones was placed in a family Cyphornithidae, and considered close allies of the pelican family (Pelicanidae). They were united with the latter in a superfamily Pelecanides in suborder Pelecanae, or later on (after the endings of taxonomic ranks were fixed to today's standard) Pelecanoidea in suborder Pelecani. Subsequently, some allied them with the entirely spurious "family" "Cladornithidae" in a "pelecaniform" suborder "Cladornithes". Those genera known from skull material were typically assigned to one or two families (Odontopterygidae and sometimes also Pseudodontornithidae) in a "pelecaniform" suborder Odontopteryges or Odontopterygia. Pelagornis meanwhile, described from wing bones, was traditionally placed in a monotypic "pelecaniform" family Pelagornithidae. This was often assigned either to the gannet and cormorant suborder Sulae (which was formerly treated as superfamily Sulides in suborder Pelecanae), or to the Odontopterygia. The sternum of Gigantornis was placed in the albatross family (Diomedeidae) in the order of tube-nosed seabirds (Procellariiformes).

The most extensive taxonomic and systematic confusion affected Dasornis. That genus was established based on a huge skull piece, which for long was placed in the Gastornithidae merely due to its size. Argillornis – nowadays recognized to belong in Dasornis – was described from wing bones, and generally included in the Sulae as part of the "Elopterygidae" – yet another invalid "family", and its type genus is generally not considered a modern-type bird by current authors. Some additional tarsometatarsus (ankle) bone fragments were placed in the genus Neptuniavis and assigned to the Procellariidae in the Procellariiformes. All these remains were only shown to belong in the pseudotooth bird genus Dasornis in 2008.