George R. Price

George R. Price

George Robert Price (October 6, 1922 – January 6, 1975) was an American population geneticist. Originally a physical chemist and later a science journalist, he moved to London in 1967, where he worked in theoretical biology at the Galton Laboratory, making three important contributions: first, rederiving W.D. Hamilton's work on kin selection with a new Price equation; second, introducing (with John Maynard Smith) the concept of the evolutionarily stable strategy (ESS), a central concept in game theory; and third, formalising Fisher's fundamental theorem of natural selection. After giving all his possessions to the poor, he committed suicide.