Cast
The fourth season had twelve roles receiving star billing. All eleven starring actors from the previous season returned for this season. The series is narrated by Brenda Strong, who portrays the deceased Mary Alice Young, as she observes from beyond the grave the lives of the Wisteria Lane residents and her former best friends. Teri Hatcher portrayed Susan Delfino, who is finally married after three seasons searching for a stable relationship. Felicity Huffman portrayed Lynette Scavo, who fights cancer during the first half of the season and deals with her step-daughter's behavior in the second half. Marcia Cross portrayed Bree Hodge, married for the second time and faking pregnancy in an attempt to protect her daughter. Eva Longoria portrayed former model Gabrielle Lang, now the wife of Fairview's mayor. Nicollette Sheridan portrayed Edie Britt, who faked suicide in the previous season finale. Ricardo Antonio Chavira portrayed Carlos Solis, now divorced from Gabrielle but still in love with her while involved in a relationship with Edie. Andrea Bowen portrayed Julie Mayer, the responsible and caring daughter of Susan. Doug Savant portrayed Tom Scavo, Lynette's husband and now owner of a pizzeria. Kyle MacLachlan portrayed Orson Hodge, Bree's second husband and the man that ran over Mike in the second season finale. James Denton portrayed Mike Delfino, now Susan's husband who starts using drugs. Dana Delany is introduced in the role of Katherine Mayfair, whose mysterious arc is the season's main storyline.
Five out of six actors who received "also starring" billing from the previous season returned. Shawn Pyfrom, portrayed Bree's homosexual son Andrew Van de Kamp. Lyndsy Fonseca was cast in the new role of Dylan Mayfair, Katherine's daughter and part of the season's mystery. Joy Lauren reappeared in the role of Danielle Van de Kamp, Bree's pregnant daughter. Brent Kinsman, Shane Kinsman and Zane Huett, played Preston Scavo, Porter Scavo and Parker Scavo, Lynette's troublesome children. Rachel Fox was promoted to regular after guest starring in several episodes in the previous season as Kayla Huntington Scavo, Tom's mischievous daughter and the result of an affair he had before meeting Lynette.
This season featured many established and new guest stars. Kathryn Joosten portrayed Karen McCluskey, one of the most prominent residents of Wisteria Lane, Richard Burgi appeared as Karl Mayer, Susan's ex-husband who is also married again, Pat Crawford Brown played elderly neighbor Ida Greenberg, whereas Tuc Watkins and Kevin Rahm made their debut in this season respectively as Bob Hunter and Lee McDermott, a gay couple from Chicago who move to the lane. Part of Lynette's storyline were Polly Bergen in the role of Stella Wingfield, Lynette's mother who moves to the Scavo family home to support her daughter in her battle against cancer, and Jason Gedrick playing Rick Coletti, who had feelings for Lynette in the previous season and now opens his own restaurant to compete with the Scavo Pizzeria. Part of Bree's storyline were Shirley Knight returning as Phyllis Van de Kamp, Bree's first mother-in-law, and Dakin Matthews appearing as Reverend Sykes, reverend at the local Presbyterian church. Part of Gabrielle's storyline were John Slattery playing Victor Lang, Gabrielle's new husband and the mayor of Fairview, Mike Farrell in the role of Milton Lang, Victor's father, Jeff Doucette portraying Father Crowley, priest at the local Catholic church, Justine Bateman playing Ellie Leonard, a drug dealer who rents a room in the Solises' house, and Jesse Metcalfe reappearing as John Rowland, Gabrielle's ex-lover and former gardener. Part of the main mystery arc were Nathan Fillion portraying Adam Mayfair, a doctor and Katherine's second husband, Ellen Geer appearing as Lillian Simms, Katherine's aunt, Melora Walters playing Sylvia Greene, Adam's former patient and ex-lover from Chicago, and Gary Cole in the role of Wayne Davis, a cop and Katherine's abusive first husband. Additionally, Gale Harold was introduced as Susan's new lover Jackson Braddock during the five-year jump featured at the end of the season.
For the 14th Annual Screen Actors Guild Awards, Fillion, Joosten and Slattery were nominated along with the rest of the cast for the Outstanding Performance by an Ensemble in a Comedy Series award due to their many appearances this season.
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