Bird Migration - Long-distance Migration

Long-distance Migration

The typical image of migration is of northern landbirds, such as swallows and birds of prey, making long flights to the tropics. However, many Holarctic wildfowl and finch species winter in the North Temperate Zone, but in regions with milder winters than their summer breeding grounds. For example, the pink-footed goose migrates from Iceland to Britain and neighbouring countries, whilst the Dark-Eyed Junco migrates from subarctic and arctic climates to the contiguous United States and the American Goldfinch from taiga to wintering grounds extending from the American South northwestward to Western Oregon. Migratory routes and wintering grounds are traditional and learned by young during their first migration with their parents. Some ducks, such as the Garganey, move completely or partially into the tropics.

The same considerations about barriers and detours that apply to long-distance land-bird migration apply to water birds, but in reverse: a large area of land without bodies of water that offer feeding sites is a barrier to may also be a barrier to a bird that feeds in coastal waters. Detours avoiding such barriers are observed: for example, Brent Geese migrating from the Taymyr Peninsula to the Wadden Sea travel via the White Sea coast and the Baltic Sea rather than directly across the Arctic Ocean and northern Scandinavia.

A similar situation occurs with waders (called "shorebirds" in North America). Many species, such as Dunlin and Western Sandpiper, undertake long movements from their Arctic breeding grounds to warmer locations in the same hemisphere, but others such as Semipalmated Sandpiper travel longer distances to the tropics in the Southern Hemisphere. Like the large and powerful wildfowl, the waders are strong fliers. This means that birds wintering in temperate regions have the capacity to make further shorter movements in the event of particularly inclement weather.

For some species of waders, migration success depends on the availability of certain key food resources at stopover points along the migration route. This gives the migrants an opportunity to "refuel" for the next leg of the voyage. Some examples of important stopover locations are the Bay of Fundy and Delaware Bay.

Some Bar-tailed Godwits have the longest known non-stop flight of any migrant, flying 11,000 km from Alaska to their New Zealand non-breeding areas. Prior to migration, 55 percent of their bodyweight is stored fat to fuel this uninterrupted journey.

Seabird migration is similar in pattern to those of the waders and waterfowl. Some, such as the Black Guillemot and some gulls, are quite sedentary; others, such as most terns and auks breeding in the temperate northern hemisphere, move varying distances south in the northern winter. The Arctic Tern has the longest-distance migration of any bird, and sees more daylight than any other, moving from its Arctic breeding grounds to the Antarctic non-breeding areas. One Arctic Tern, ringed (banded) as a chick on the Farne Islands off the British east coast, reached Melbourne, Australia in just three months from fledging, a sea journey of over 22,000 km (14,000 mi). A few seabirds, such as Wilson's Petrel and Great Shearwater, breed in the southern hemisphere and migrate north in the southern winter. Seabirds have the additional advantage of being able to feed during migration over open waters.

The most pelagic species, mainly in the 'tubenose' order Procellariiformes, are great wanderers, and the albatrosses of the southern oceans may circle the globe as they ride the "roaring forties" outside the breeding season. The tubenoses spread widely over large areas of open ocean, but congregate when food becomes available. Many are also among the longest-distance migrants; Sooty Shearwaters nesting on the Falkland Islands migrate 14,000 km (8,700 mi) between the breeding colony and the North Atlantic Ocean off Norway. Some Manx Shearwaters do this same journey in reverse. As they are long-lived birds, they may cover enormous distances during their lives; one record-breaking Manx Shearwater is calculated to have flown 8 million km (5 million miles) during its over-50 year lifespan.

Some large broad-winged birds rely on thermal columns of rising hot air to enable them to soar. These include many birds of prey such as vultures, eagles, and buzzards, but also storks. These birds migrate in the daytime. Migratory species in these groups have great difficulty crossing large bodies of water, since thermals only form over land, and these birds cannot maintain active flight for long distances. The Mediterranean and other seas present a major obstacle to soaring birds, which must cross at the narrowest points. Massive numbers of large raptors and storks pass through areas such as Gibraltar, Falsterbo, and the Bosphorus at migration times. More common species, such as the European Honey Buzzard, can be counted in hundreds of thousands in autumn. Other barriers, such as mountain ranges, can also cause funnelling, particularly of large diurnal migrants. This is a notable factor in the Central American migratory bottleneck. Batumi bottleneck in the Caucasus is one of the heaviest migratory funnels on earth. Avoiding flying over the Black Sea surface and across high mountains, hundreds of thousands of soaring birds funnel through an area around the city of Batumi, Georgia.

Many of the smaller insectivorous birds including the warblers, hummingbirds and flycatchers migrate large distances, usually at night. They land in the morning and may feed for a few days before resuming their migration. The birds are referred to as passage migrants in the regions where they occur for short durations between the origin and destination.

Nocturnal migrants minimize predation, avoid overheating, and feed during the day. One cost of nocturnal migration is the loss of sleep. Migrants may be able to alter their quality of sleep to compensate for the loss.

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