Fossil Record
The fossil record of treeshrews is poor. The oldest putative treeshrew, Eodendrogale parva, is from the Middle Eocene of Henan, China, but the identity of this animal is uncertain. Other fossils have come from the Miocene of Thailand, Pakistan, India, and Yunnan, China, as well as the Pliocene of India. Most belong to the family Tupaiidae, but some still-undescribed fossils from Yunnan are thought to be closer to the pen-tailed treeshrew (Ptilocercus). Named fossil species include Prodendrogale yunnanica, Prodendrogale engesseri, and Tupaia storchi from Yunnan, Tupaia miocenica from Thailand, and Palaeotupaia sivalicus from India.
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