Tortoise - Taxonomy

Taxonomy

The following species list largely follows Rhodin et al., 2010, this is a work in progress.

Family Testudinidae Batsch 1788

Subfamily Testudininae

• Aldabrachelys
  • Aldabrachelys gigantea, Aldabra giant tortoise, common synonyms Geochelone gigantea, Dipsochelys gigantea
  • †Aldabrachelys abrupta (extinct)

• Astrochelys Gray, 1873
  • Astrochelys radiata, radiated tortoise
  • Astrochelys yniphora, angulated tortoise, (Madagascan) plowshare tortoise, angonoka

• Chelonoidis Fitzinger, 1835.
  • Chelonoidis carbonaria, red-footed tortoise
  • Chelonoidis chilensis, pampas tortoise, South Argentine tortoise
  • Chelonoidis denticulata, Brazilian giant tortoise, yellow-footed tortoise
  • Chelonoidis nigra complex:
    • †Chelonoidis abingdonii, Pinta Island giant tortoise, Abingdon Island giant tortoise (extinct)
    • Chelonoidis becki, Volcan Wolf giant tortoise, Cape Berkeley giant tortoise
    • Chelonoidis chathamensis, San Cristobal giant tortoise, Chatham Island giant tortoise
    • Chelonoidis darwini, San Salvador giant tortoise, James Island giant tortoise
    • Chelonoidis duncanensis, Pinzon giant tortoise, Duncan Island giant tortoise
    • Chelonoidis hoodensis, Espanola giant tortoise, Hood Island giant tortoise
    • †Chelonoidis nigra, Floreana giant tortoise, Charles Island giant tortoise (extinct)
    • †Chelonoidis phantastica, Fernandina giant tortoise, Narborough Island giant tortoise (extinct)
    • Chelonoidis porteri, Santa Cruz giant tortoise, Indefatigable Island giant tortoise
    • Chelonoidis vicina, Isabela Island giant tortoise, Albemarle Island giant tortoise
  • Chelonoidis petersi, Chaco tortoise, northern Argentine tortoise

• Chersina
  • Chersina angulata, Angulate Tortoise, South African bowsprit tortoise

• Colossochelys
  • †Colossochelys atlas, Atlas tortoise (extinct)

• †Cylindraspis (all species extinct) following Austin and Arnold, 2001:
  • Cylindraspis indica, synonym Cylindraspis borbonica, Reunion giant tortoise
  • Cylindraspis inepta, Mauritius giant domed tortoise
  • Cylindraspis peltastes, Rodrigues domed tortoise
  • Cylindraspis triserrata, Mauritius giant flat-shelled tortoise
  • Cylindraspis vosmaeri, Rodrigues giant saddleback tortoise

• Geochelone
  • Geochelone elegans, Indian star tortoise
  • Geochelone platynota, Burmese star tortoise
  • Geochelone sulcata, African spurred tortoise (Sulcata tortoise)

• †Hadrianus
  • Hadrianus corsoni (syn. H. octonarius)
  • Hadrianus robustus
  • Hadrianus schucherti
  • Hadrianus utahensis

• Homopus
  • Homopus areolatus, parrot-beaked tortoise, beaked Cape tortoise, common padloper
  • Homopus boulengeri, Karoo dwarf tortoise, Boulenger's Cape tortoise, Karoo padloper
  • Homopus femoralis, greater dwarf tortoise, greater padloper
  • Homopus signatus, speckled tortoise, speckled padloper, speckled padloper tortoise
  • Homopus solus, Nama tortoise, Berger's cape tortoise, Nama padloper, synonym Homopus bergeri

• Indotestudo
  • Indotestudo elongata, elongated tortoise, yellow-headed tortoise
  • Indotestudo forstenii, Forsten’s tortoise, East Indian tortoise
  • Indotestudo travancorica, Travancore tortoise

• Kinixys
  • Kinixys belliana, Bell's hinge-back tortoise
  • Kinixys erosa, Forest hinge-back tortoise, serrated hinge-back tortoise
  • Kinixys homeana, Home's hinge-back tortoise
  • Kinixys lobatsiana, Lobatse hinge-back tortoise
  • Kinixys natalensis, Natal hinge-back tortoise
  • Kinixys spekii, Speke's hinge-back tortoise

• Malacochersus
  • Malacochersus tornieri, pancake tortoise

• Psammobates
  • Psammobates geometricus, geometric tortoise
  • Psammobates oculifer, serrated tent tortoise, Kalahari tent tortoise
  • Psammobates tentorius, African tent tortoise

• Pyxis
  • Pyxis arachnoides, (Madagascan) spider tortoise
  • Pyxis planicauda, flat-backed spider tortoise, (Madagascan) flat-tailed tortoise, flat-tailed spider tortoise

• Stigmochelys Gray, 1873
  • Stigmochelys pardalis, leopard tortoise

• Testudo
  • Testudo graeca, Greek tortoise, spur-thighed tortoise, Moorish tortoise
  • Testudo hermanni, Hermann's tortoise
  • Testudo horsfieldii, Russian tortoise, steppe tortoise, Horsfield's tortoise, or Central Asian tortoise
  • Testudo kleinmanni, Egyptian tortoise, including Negev tortoise
  • Testudo marginata, marginated tortoise

Subfamily Xerobatinae Agassiz, L. 1857.

• Gopherus
  • Gopherus agassizii, Mojave Desert tortoise, Agassiz's desert tortoise
  • Gopherus berlandieri, Texas tortoise, Berlandier's tortoise
  • Gopherus flavomarginatus, Bolson tortoise
  • Gopherus morafkai, Sonoran Desert tortoise, Morajak's desert tortoise
  • Gopherus polyphemus, gopher tortoise

• Manouria
  • Manouria emys, Asian giant tortoise, brown tortoise (mountain tortoise)
  • Manouria impressa, impressed tortoise

• †Stylemys (Genus extinct)
  • Stylemys botti
  • Stylemys calaverensis
  • Stylemys canetotiana
  • Stylemys capax
  • Stylemys conspecta
  • Stylemys copei
  • Stylemys emiliae
  • Stylemys frizaciana
  • Stylemys karakolensis
  • Stylemys nebrascensis (syn. S. amphithorax)
  • Stylemys neglectus
  • Stylemys oregonensis
  • Stylemys pygmea
  • Stylemys uintensis
  • Stylemys undabuna