Cast
- Gaynor Faye as Anna Williams - the daughter of the late Imogen and Frank but she was brought up by George and Imogen Williams and sister to Sarah, cousin to Fiona and wife to tom. Mother to Daisy, Luca, Jake and baby Imogen And a very experienced vet.
- Nicola Stephenson as Sarah Williams - the daughter of the late Imogen and only biological daughter of George Williams and sister to Anna, Sarah moved to London following the death of her mother and the discovery of Tom's affair with Fiona. Having returned to Oxley, Sarah's a confident vet. Sarah marries Adrian and becomes pregnant, but loses the baby. After discovering that her sister and husband discussed her fertility problems behind her back, she sleeps with Sebastian. She laters finds out that she is pregnant with his child.
- Nicholas Gleaves as Tom Bedford - Husband to Anna and father to Daisy, Luca, Jake, baby Imogen and Theo Jones. He had a one night stand with Fiona, which resulted in her getting pregnant with Theo.
- Heather Peace as Fiona Jones - single mother to Theo, the result of a drunken one-night stand with Tom. She has her heart set on getting Tom for herself and Theo. Brought up by a single mum, Fiona's like the third sister of the Williams clan and constantly feels like Cinderella. She was the one in the hand-me-down clothes, who trained to be a nurse not a vet, and who couldn't get the man.
- Michelle Holmes as Claudie Williams - widow of George. She was genuinely in love with him but also conscious that he was sitting on a gold mine. She happily encouraged him to re-mortgage the house to fund their lengthy honeymoon stay in Malta.
- Janet Dibley as Deborah Johnson - Rick's domineering wife Deborah is the very bossy Practice Manager of The Chase. An old nag, and snobby to go with it, Deborah doesn't have a nice word to say about anyone or anything. She's always right and happy to let everyone know.
- Reece Dinsdale as Rick Johnson - The hen-pecked husband of Deborah, Rick's a Senior Partner at The Chase. He's put upon and nagged to death by his wife but he loves his work and is a close associate of the Williams family. The only person who really seems to understand what he goes through is Clare.
- Rachel Davies as Margaret Wright - A Practice Nurse, Margaret's been at The Chase since the building was built. Margaret is always there - and always happy to gossip about it. Although undoubtedly good at her job, she's very scatty and liable to get stressed at the drop of a hat.
- Sunetra Sarker as Clare Burnes - Clare is the Practice Nurse at The Chase. Her rip-roaring sense of humour keeps everyone happy, especially Rick.
- Liam Garrigan as Matt Lowe - The most junior Practice Nurse at The Chase, Matt has dreams of leaving it all behind and becoming a rock star. He had a short relationship with Anna.
- Paul Opacic as Adrian Huby - local farmer and Sarah's first love, who marries Sarah in series 2.
- Rebekah Manning as Paula Higgins - Paula is Claudie's younger sister. Claudie set Paula up as a beautician in order to turn her life around. Paula arrives in the village to be with her sister and causes chaos in the practice as half the contents of her salon seem to arrive with her. Blonde and bubbly, she is close to her sister but likes to remain on good terms with everyone.
- Christian Cooke as Liam Higgins - Liam is Claudie and Paula's younger brother. Only nineteen and a bit of a rogue, he appears during the series after having been let out of a young offenders institute.
- Philip Jackson as Frank Jones - Fiona's estranged father, Frank returns to Oxley unexpectedly. He doesn't have many friends in the village, as it is common knowledge that he left his wife and young daughter for another woman. Only Fiona knows that the other woman was in fact Anna and Sarah's mother Imogen, who decided to remain with George and the girls at the last minute. He is secretly Anna's father.
- Guy Henry as William Montgomery - The eldest son of the late Lord Montgomery, William has taken his father's title and responsibility of the estate. The Chase's best client, he holds a long standing respect for George and his two daughters.
- Adam Croasdell as Sebastian Montgomery - William Montgomery's younger brother and old friend of Adrian Huby. Sebastian is good looking and charming, and keen to avoid any of the business of the family estate. Involved in the music business, he is good at making money, and attracting female attention.
- Grace Cassidy as Harriet Johnson - Rick and Deborah's 15 year old daughter - as in series 2
- Grace Ruiz as Daisy Williams
- Nathan Chaplow as Jake Williams
- Eliot Anthony as Luca Williams
- Charlie Connew as Theo Jones
- Keith Barron as George Williams (Series 1) - George was the father of Anna and Sarah and the head vet of The Chase. His first wife, Imogen, died. Recently, he married Claudie. George was much respected in the local community for always being available to treat farm animals, even on Christmas Day. His principles meant that the practice was somewhat strapped for cash though. Fiona murdered him by cutting the brakes in the Land Rover - however, she had intended to kill Sarah, and was devastated when her "uncle", whom she truly loved, died instead.
- Some notes are taken from .
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