Dominance Hierarchy
Space inside the nest may also be divided as a result of dominance interactions. For example, in paper wasp colonies, a single inseminated queen may found (initiate) a colony after waking up from hibernation (overwintering). However, it is common in many species that multiple inseminated females join these foundresses instead of founding their own nest. When multiple inseminated females found a colony together, the colony grows quickly, yet only one individual will become the primary egg-layer. Through a series of dominance interactions, the most aggressive wasp will emerge as the dominant individual and will become the primary egg-layer for the group (the prime role for ensuring your genes are passed on to subsequent generations), whereas the remaining subordinate wasps will perform other tasks, such as nest construction or foraging. There is evidence that these dominance interactions affect the spatial zones individuals occupy as well. In paper wasps (Ropalidia revolutionalis), as well as in the ant species Odontomachus brunneus, dominant individuals are more likely to reside in the central areas of the nest, where they take care of brood, while the subordinate individuals are pushed towards the edge, where they are more likely to forage. It is unknown whether division of space or establishment of dominance occurs first and if the other is a result of it.
Read more about this topic: Spatial Organization, Spatial Organization in Eusocial Insects, Spatial Organization in The Nest
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