Scientific Contributions
Robert Darwin provided the first empirical evidence that small eye movements are made even when people attempt to keep them fixed. This he found during his studies of the afterimages of colored stimuli in which he noticed that while a person tried to fixate a colored circle, a lucid edge appeared on the adjacent white-paper background. He concluded "as by the unsteadiness of the eye a part of the fatigued retina falls on the white paper".
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