Early Life and Education
Howard was born in Stamford, Connecticut and raised in Warren, Ohio. He received an A.B. from Miami University in 1938 and took up a position as a technician with Irving W. Bailey at Harvard University. Howard worked on the Icacinaceae plant family and in 1939 he received a fellowship which supported his graduate study. He continued to work on the Icacinaceae, receiving his doctorate in 1942. His doctoral thesis was entitled "Studies of the Icacinaceae: A monograph of the New World species.
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