Process
When a molecule of chlorophyll absorbs light it is promoted from its ground state to its first singlet excited state. The excited state then has three main fates. Either the energy is; 1. passed to another chlorophyll molecule by Förster resonance energy transfer (in this way excitation is gradually passed to the photochemical reaction centers (photosystem I and photosystem II) where energy is used in photosynthesis (called photochemical quenching); or 2. the excited state can return to the ground state by emitting the energy as heat (called non-photochemical quenching); or 3. the excited state can return to the ground state by emitting a photon (fluorescence).
In higher plants, the absorption of light continues to increase as light intensity increases, while the capacity for photosynthesis tends to saturate. Therefore there is the potential for the absorption of excess light energy by photosynthetic light harvesting systems. This excess excitation energy leads to an increase in the lifetime of singlet excited chlorophyll, increasing the chances of the formation of long-lived chlorophyll triplet states by inter-system crossing. Triplet chlorophyll is a potent photosensitiser of molecular oxygen forming singlet oxygen which can cause oxidative damage to the pigments, lipids and proteins of the photosynthetic thylakoid membrane. To counter this problem, one photoprotective mechanism is so-called non-photochemical quenching (NPQ), which relies upon the conversion and dissipation of the excess excitation energy into heat. NPQ involves conformational changes within the light harvesting proteins of photosystem II that bring about a change in pigment interactions causing the formation of energy traps. The conformational changes are stimulated by a combination of transmembrane proton gradient, the PsbS subunit of photosystem II and the enzymatic conversion of the carotenoid violaxanthin to zeaxanthin (the xanthophyll cycle).
For further discussion, see Chlorophyll fluorescence and Plant stress measurement.
Read more about this topic: Non-photochemical Quenching
Famous quotes containing the word process:
“It is part of the nature of consciousness, of how the mental apparatus works, that free reason is only a very occasional function of people’s “thinking” and that much of the process is made of reactions as standardized as those of the keys on a typewriter.”
—John Dos Passos (1896–1970)
“The process of education in the oldest profession in the world is like any other educational process, in that it requires time and effort and patience; it can only be acquired by taking one step at a time, though the steps become accelerated after the first few.”
—Madeleine [Blair], U.S. prostitute and “madam.” Madeleine, ch. 4 (1919)
“The moralist and the revolutionary are constantly undermining one another. Marx exploded a hundred tons of dynamite beneath the moralist position, and we are still living in the echo of that tremendous crash. But already, somewhere or other, the sappers are at work and fresh dynamite is being tamped in place to blow Marx at the moon. Then Marx, or somebody like him, will come back with yet more dynamite, and so the process continues, to an end we cannot foresee.”
—George Orwell (1903–1950)