Mollusca - Definition

Definition

The two most universal features of the body structure of molluscs are a mantle with a significant cavity used for breathing and excretion, and the organization of the nervous system. The most abundant metallic element in molluscs is calcium.

Molluscs have developed such a varied range of body structures, it is difficult to find synapomorphies (defining characteristics) to apply to all modern groups. The most general characteristic of molluscs is they are unsegmented and bilaterally symmetrical. The following are present in all modern molluscs:

  • The dorsal part of the body wall is a mantle (or pallium) which secretes calcareous spicules, plates or shells. It overlaps the body with enough spare room to form a mantle cavity.
  • The anus and genitals open into the mantle cavity.
  • There are two pairs of main nerve cords.

Other characteristics that commonly appear in textbooks have significant exceptions:

Class
Characteristic Aplacophora Polyplacophora Monoplacophora Gastropoda Cephalopoda Bivalvia Scaphopoda
Radula, a rasping "tongue" with chitinous teeth Absent in 20% of Neomeniomorpha Yes Yes Yes Yes No Internal, cannot extend beyond body
Broad, muscular foot Reduced or absent Yes Yes Yes Modified into arms Yes Small, only at "front" end
Dorsal concentration of internal organs (visceral mass) Not obvious Yes Yes Yes Yes Yes Yes
Large digestive ceca No ceca in some aplacophora Yes Yes Yes Yes Yes No
Large complex metanephridia ("kidneys") None Yes Yes Yes Yes Yes Small, simple

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