Evolution
The Theolinae evolved in a tropical climate while the Lycaenidae and Icaricia evolved in a temperate climate zone. The species Icaricia icaroides comprises 12 subspecies. They are, I. icarioides, I. evius, I. moroensis, I. missionensis, I. padalis, I. pheres, I. ardea, I. lycea, I. bucholzi, I. pembina, I. blackmorei, and I. montis. The genus Icaricia was introduced in a 1944 paper by Vladimir Nabokov. The paper, Notes on the morphology of the genus Lycæides (Lycænidæ, Lepidoptera), was published in the Sep.-Dec. issue of Psyche—A Journal of Entomology and described two new genera, Icaricia and Plebulina.
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