Breeding
They breed in North America from as far east as Quebec to the Pacific coast and Alaska, and from Finland and Estonia across northern Asia. They are permanent residents, but may move south and southeast when food is scarce. A small population, estimated at less than 100 birds, occurs in the Sierra Nevada mountains of California. This population is the southernmost population of the species' range and is listed Endangered under California's Endangered Species Act.
Their breeding habitat is the dense coniferous forests of the taiga, near open areas, such as meadows or bogs. Great Grey Owls do not build nests, so typically use nests previously used by a large bird, such as a raptor. They will also nest in broken-topped trees and cavities in large trees. Nesting may occur from March to May. Four eggs are the usual clutch size. Eggs average 42.7 mm wide and 53.5 mm long (1.68 by 2.11 in). The incubation period is about 30 days, ranging from 28 to 36 days. Brooding lasts 2 to 3 weeks, after which the female starts roosting on a tree near nests. The young jump or fall from the nest at 3 to 4 weeks, and start to fly 1 to 2 weeks after this. Most offspring remain near their natal sites for many months after fledging.
The abundance of food in the area usually affects the number of eggs a female lays, a feature quite common in northern owl species. If food is scarce, they may travel a fair distance to find more prey, with considerable movements by large numbers in some years of particularly scarce prey. Though they do not migrate, many are at least somewhat nomadic.
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