Banksia Ser. Abietinae - According To Thiele and Ladiges

According To Thiele and Ladiges

In 1996, Kevin Thiele and Pauline Ladiges undertook a cladistic analysis of morphological characters of Banksia, which yielded a phylogeny somewhat at odds with George's taxonomic arrangement. Their cladogram included a clade consisting of the members of B. ser. Abietinae sensu George, and also B. tricuspis (Lesueur Banksia):

Thiele and Ladiges therefore retained George's B. ser. Abietinae largely intact, but transferred Banksia tricuspis (Lesueur Banksia) back into it. They then further divided it into four subseries, in accordance with the resolution of clades in their analysis. The placement and circumscription of B. ser. Abietinae in Thiele and Ladiges' arrangement may be summarised as follows:

Banksia

B. subg. Isostylis (3 species)

B. elegans (incertae sedis)

B. subg. Banksia

B. ser. Tetragonae (4 species)

B. ser. Lindleyanae (1 species)

B. ser. Banksia (2 subseries, 12 species)

B. baueri (incertae sedis)

B. lullfitzii (incertae sedis)

B. attenuata (incertae sedis)

B. ashbyi (incertae sedis)

B. coccinea (incertae sedis)

B. ser. Prostratae (8 species)

B. ser. Cyrtostylis (4 species)

B. ser. Ochraceae (3 species, 2 subspecies)

B. ser. Grandes (2 species)

B. ser. Salicinae (2 subseries, 11 species, 4 subspecies)

B. ser. Spicigerae (3 subseries, 7 species, 6 varieties)

B. ser. Quercinae (2 species)

B. ser. Dryandroideae (1 species)

Banksia ser. Abietinae

Banksia subser. Nutantes

B. nutans

B. n. var. nutans

B. n. var. cernuella

Banksia subser. Sphaerocarpae

B. grossa

B. dolichostyla (now B. sphaerocarpa var. dolichostyla)

B. micrantha

B. sphaerocarpa

B. s. var. sphaerocarpa

B. s. var. caesia

Banksia subser. Leptophyllae

B. telmatiaea

B. scabrella

B. leptophylla

B. l. var. melletica

B. l. var. leptophylla

B. lanata

Banksia subser. Longistyles

B. violacea

B. laricina

B. incana

B. tricuspis

B. pulchella

B. meisneri

B. m. var. meisneri

B. m. var. ascendens

Thiele and Ladiges' arrangement remained current only until 1999, when George's treatment of the genus for the Flora of Australia series of monographs was published. This was essentially a revision of George's 1981 arrangement, which took into account some of Thiele and Ladiges' data, but rejected their overall arrangement. With respect to B. ser. Abietinae, George's 1999 arrangement differed from his 1981 arrangement only in the ordering of the species.