Ammonoidea - Distribution

Distribution

Starting from the mid-Devonian, ammonoids were extremely abundant, especially as ammonites during the Mesozoic era. Many genera evolved and ran their course quickly, becoming extinct in a few million years. Due to their rapid evolution and widespread distribution, ammonoids are used by geologists and paleontologists for biostratigraphy. They are excellent index fossils, and it is often possible to link the rock layer in which they are found to specific geological time periods.

Due to their free-swimming and/or free-floating habits, ammonites often happened to live directly above seafloor waters so poor in oxygen as to prevent the establishment of animal life on the seafloor. When upon death the ammonites fell to this seafloor and were gradually buried in accumulating sediment, bacterial decomposition of these corpses often tipped the delicate balance of local redox conditions sufficiently to lower the local solubility of minerals dissolved in the seawater, notably phosphates and carbonates. The resulting spontaneous concentric precipitation of minerals around a fossil, a concretion, is responsible for the outstanding preservation of many ammonite fossils.

When ammonites are found in clays, their original mother-of-pearl coating is often preserved. This type of preservation is found in ammonites such as Hoplites from the Cretaceous Gault clay of Folkestone in Kent, England.

The Cretaceous Pierre Shale formation of the United States and Canada is well known for the abundant ammonite fauna it yields, including Baculites, Placenticeras, Scaphites, Hoploscaphites, and Jeletzkytes, as well as many uncoiled forms. Many of these also have much or all of the original shell, as well as the complete body chamber, still intact. Many Pierre Shale ammonites, and indeed many ammonites throughout earth history, are found inside concretions.

Other fossils, such as many found in Madagascar and Alberta, display iridescence. These iridescent ammonites are often of gem quality (ammolite) when polished. In no case would this iridescence have been visible during the animal's life; additional shell layers covered it.

The majority of ammonoid specimens, especially those of the Paleozoic era, are preserved only as internal molds; the outer shell (composed of aragonite) has been lost during the fossilization process. Only in these internal-mold specimens can the suture lines be observed; in life, the sutures would have been hidden by the outer shell.

The ammonoids as a group continued through several major extinction events, although often only a few species survived. Each time, however, this handful of species diversified into a multitude of forms. Ammonite fossils became less abundant during the latter part of the Mesozoic, with none surviving into the Cenozoic era. The last surviving lineages disappeared, along with the dinosaurs, 65 Mya in the Cretaceous–Paleogene extinction event. The reason why no ammonites survived the extinction event at the end of the Cretaceous, whereas some nautiloids survived, might be due to differences in ontogeny. If their extinction was due to a bolide strike, plankton around the globe could have been severely diminished, thereby dooming ammonite reproduction during its planktonic stage.